Vegetation of the eastern communal conservancies in Namibia: II. Environmental drivers

The eastern communal conservancies are situated along the western fringe of the Kalahari basin. Under a very short rainfall gradient, the vegetation abruptly changes from microphyllous Acacia-dominated savannas to mesophyll savannas, dominated by Terminalia sericea and Combretum spp. We hypothesise that this is caused by changes in soil moisture availability brought about by changes in soil texture from loamy soils to deep sands (the ‘inverse texture effect’). For this analysis, we used vegetation and soils data derived from a recognisance survey of the natural resources of the study area. As the sites in the soil and vegetation surveys did not overlap, it was decided to use only synoptic data for the plant associations in the analysis. Non-metric multidimesional scaling ordination was utilised as ordination technique of the vegetation data and various environmental parameters, including soil texture, soil hydraulic parameters, climatic and fire regime parameters, were overlaid as biplots onto the resulting graph, as were various plant functional attributes particularly related to climatic conditions. The main environmental gradient identified within the study area is the rainfall gradient. This relatively short gradient, however, does not explain the marked change in vegetation observed within the study area. This change is attributed to the change in soil type, in particular, the soil texture and the associated soil hydraulic parameters of the soil. This gradient is closely correlated to leaf size, explaining the change from microphyll savannas to mesophyll savannas along the change from loamy to sandy soils. One of the lesser understood mechanisms for the survival of these mesophyll plants on sandy soils seems to be a deep root system, which is actively involved in water redistribution within the soil profile – by hydraulic lift, inverse hydraulic lift and stem flow.